• Alburquerque, J.A. Salazar, P., Barrón, V., Torrent, J. Campillo, M.C., Gallardo, A., Villar, R. 2013. Enhanced wheat yield by biochar addition under different mineral fertilization levels. Agron. Sustain. Dev. Pdf


  • Alameda, D. Anten, N., Villar, R. 2012. Soil compaction effects on growth and root traits of tobacco depend on light, water regime and mechanical stress. Soil and Tillage Research 120: 121-129. Pdf
  • Alameda, D., Villar, R., Iriondo, J.M. 2012. Spatial pattern of soil compaction: Trees´ footprint on physical properties. Forest Ecology and Management,  283: 128–137. Pdf
  • Alameda, D., Villar, R., 2012. Linking root traits to plant physiology and growth in Fraxinus angustifolia Vahl. seedlings under soil compaction conditions. Environmental and Experimental Botany 79: 49– 57. Pdf
  • Dominguez, M. T., Aponte, C., Pérez-Ramos, I.M., García, L.V., Villar, R., Marañón, T. 2012. Relationships between leaf morphological traits, nutrient concentrations and isotopic signatures for Mediterranean woody plant species and communities. Plant and Soil 357:407–424. Pdf
  • González-Rodríguez, V., Barrio, I.C., Villar, R. 2012. Within-population variability influences early seedling establishment in four Mediterranean  oaks. Acta Oecologica 41: 82-89. Pdf
  • González-Rodríguez, V., Villar, R. 2012. Post-dispersal seed removal in four Mediterranean oaks: species and microhabitat selection differ depending on large herbivore activity. Ecological Research 27 (3): 587-594. Pdf


  • Becerra, P., González-Rodríguez, V., Smith, C., Armesto, J. 2011. Spatio-temporal variation in the effect of herbaceous layer on woody seedling survival in a Chilean mediterranean ecosystem. Journal of Vegetation Science 22: 847-855. Pdf
  • González-Rodríguez, V. 2011. Establecimiento de cuatro especies de Quercus en el sur de la Península Ibérica. Factores condicionantes. Ecosistemas 20(1): 148-154. Pdf
  • González-Rodríguez, V., Villar,R., Casado, R., Suárez-Bonet, E., Quero, J.L., Navarro-Cerrillo, RM. 2011. Spatio-temporal heterogeneity effects on seedling growth and establishment in four Quercus species. Annals of Forest Science 68: 1217-1232. Pdf
  • González-Rodríguez, V., Navarro-Cerrillo, R., Villar, R. 2011. Artificial regeneration with Quercus ilex L. and Q. suber L. oaks by direct seeding and planting in southern Spain. Annals of Forest Science 68: 637-646. Pdf
  • González-Rodríguez, V., Villar, R., Navarro-Cerrillo, R. 2011. Maternal influences on seed-mass effect and initial seedling growth in four Quercus species. Acta Oecologica 37:1-9. Pdf
  • Oliet JA, Salazar JM, Villar R, Robredo E, Valladares F. 2011. Fall fertilization of Holm oak affects N and P dynamics, root growth potential, and post-planting phenology and growth. Annals of Forest Science 68 : 647–656. Pdf
  • Quero JL., FJ. Sterck, R. Villar, J. Martínez-Vilalta 2011. Water use strategies of six co-existing Mediterranean woody species during a summer drought. Oecologia 166: 45-57. Pdf


  • Bejarano MD, Villar R, Murillo A, Quero JL. 2010. Effects of soil compaction and light on growth of Quercus pyrenaica Willd. (Fagaceae) seedlings.  Soil & Tillage Research 110: 108–114. Pdf
  • Perez-Ramos IM, Gomez-Aparicio L, Villar R, García LV, Marañón T. 2010. Seedling growth and morphology of three oak species along field resource gradients and seed mass variation: a seedling age-dependent response. Journal of Vegetation Science 21: 419–437. Pdf


  • Poorter H, Niinemets U, Poorter L, Wright IJ, Villar R. 2009. Causes and consequences of variation in leaf mass per area (LMA): a meta-analysis.New Phytologist 182: 565–588. Pdf
  • Alameda D, Villar R. 2009. Moderate soil compaction: Implications on growth and architecture in seedlings of 17 woody plant species.Soil & Tillage Research 103: 325–331. Pdf


  • Quero JL, Villar R, Marañón T, Zamora R, Vega D, Sack L. 2008. Relating leaf photosynthetic rate to whole-plant growth: drought and shade effects on seedlings of four Quercus species.Functional Plant Biology 35: 725-737. Pdf
  • Quero JL, Villar R, Marañón T, Murillo A, Zamora R. 2008. Respuesta plástica a la luz y al agua en cuatro especies mediterráneas del género Quercus (Fagaceae). Revista Chilena de Historia Natural 81: 373-385. Pdf


  • Quero JL, Villar R, Marañon T, Zamora R, Poorter L (2007). Seed mass effects in four mediterranean Quercus species (Fagaceae) growing in contrasting light environments. American Journal of Botany 94(11) 1795-1803. Pdf


  • Villar R., Ruiz-Robleto J., De Jong Y., Poorter H. (2006). Differences in construction costs and chemical composition between deciduous and evergreen woody species are small as compared to differences among families. Plant, Cell and Environment 29: 1629–1643. Pdf
  • Quero, J.L, Villar R., Marañon T., Zamora R. (2006) Interactions of drought and shade effects on seedlings of four Quercus species: physiological and structural leaf responses.New Phytologist 170: 819–834. Pdf


  • Ruíz-Robleto J, Villar R. (2005) Relative growth rate and biomass allocation in ten woody species with different leaf longevity using phyllogenetic independent contrasts (PICs).Plant Biology 7: 484-494. Pdf
  • Wright  IJ, Reich PB, Cornelissen JHC, Falster DS, Garnier E, Hikosaka K, Lamont BB, Lee W, Oleksyn J, Osada N, Poorter H, Villar R, Warton DI, Westoby M (2005) Assesing the generality of global leaf trait relationships.New Phytologist 166: 485-496. Pdf


  • Villar R,  Marañón T, Quero JL, Panadero P, Arenas F, Lambers H. 2004. Variation in growth rate of 20 Aegilops species (Poaceae) in the field: The importance of net assimilation rate or specific leaf area depends on the time scale. Plant and Soil 272: 11-27. Pdf
  • Wright IJ, Reich PB, Westoby M, Ackerly DD, Baruch Z, Bongers F, Cavender-Bares J, Chapin FS, Cornelissen JHC, Diemer M, Flexas J, Garnier E, Groom PK, Gulias J, Hikosaka K, Lamont BB, Lee T, Lee W, Lusk C, Midgley JJ, Navas M-L, Niinemets Ü, Oleksyn J, Osada N, Poorter H, Poot P, Prior L, Pyankov VI, Roumet C, Thomas SC,  Tjoelker MG, Veneklaas EJ, Villar R. 2004. The world-wide leaf economics spectrum. Nature, 428: 821-827. Pdf
  • Marañón T, Zamora R, Villar R, Zavala MA, Quero JL, Pérez-Ramos I, Mendoza I, Castro J (2004). Regeneration of tree species and restoration under contrasted Mediterranean habitats: field and glasshouse experiments. International Journal of Ecology and Environmental Sciences 30 (3); 187-196.


  • Antúnez I, Retamosa EC, Villar R. (2001). Relative growth rate in phylogenetically related deciduous and evergreen woody species.Oecologia 128: 172-180. Pdf
  • Villar R, Merino JA (2001). Comparison of leaf construction costs in woody species with differing leaf life-spans in contrasting ecosystems. New Phytologist 151: 213-226.


  • Villar R, Veneklaas EJ, Jordano P, Lambers H (1998) Relative growth rate and biomass allocation in 20 Aegilops (Poaceae) species.New Phytologist 140: 425-437. Pdf


  • Lambers H, van den Boogard R, Veneklaas EJ, Villar R (1995) Carbon partitioning in wheat cultivars and wild wheat species and Global Environmental Change. Global Change Biology 1: 397-406.
  • Villar R, Held AA, Merino J (1995) Dark leaf respiration in light and darkness of an evergreen and a deciduous plant species. Plant Physiology 107: 421-427. Pdf
  • Atkin OK, Villar R, Lambers H (1995) Partitioning of electrons between the cytochrome and alternative patways in intact roots. Plant Physiology 108: 1179-1183. Pdf


  • Villar R, Held AA, Merino J (1994) Comparison of methods to estimate dark respiration in the light in leaves of two woody species.Plant Physiology 105:167-172. Pdf


  • Atkin OK, Villar R, Cummins WR (1993) The ability of several high arctic plant species to utilize nitrate nitrogen under field conditions.Oecologia 96: 239-245. Pdf


  • Collier DE, Cummins WR, Villar R (1992) Diurnal patterns of respiration in the leaves of four forest tree species.Physiologia Plantarum 84: 361-366.